[1] Univ Sains Malaysia, Ctr Marine & Coastal Studies CEMACS, Gelugor 11800, Malaysia
[2] Natl Marine Environm Monitoring Ctr, 42 Linghe St, Dalian 116023, Peoples R China
[3] Univ Sains Malaysia, Sch Humanities, Geog Sect, Gelugor 11800, Malaysia
[4] Penang Inst, Heritage & Urban Studies, Georgetown 10350, Malaysia
[5] Univ Sains Malaysia, Sch Biol Sci, Gelugor 11800, Malaysia