[1] Univ Florida, Dept Biol, Gainesville, FL 32611 USA
[2] Univ Sunshine Coast, Trop Forests & People Res Ctr, Maroochydore, Qld 4558, Australia
[3] Atlanta Bot Garden, 1345 Piedmont Ave NE, Atlanta, GA 30309 USA
[4] Univ Florida, Sch Forest Fisheries & Geomatics Sci, Gainesville, FL 32611 USA