[1] Miami Univ, Project Dragonfly, Dept Biol, Oxford, OH 45056 USA
[2] Univ Texas Austin, Primate Mol Ecol & Evolut Lab, Dept Anthropol, Austin, TX 78712 USA
[3] Miami Univ, Dept Anthropol, Oxford, OH 45056 USA
[4] Univ San Francisco Quito, Coll Biol & Environm Sci, Quito, Ecuador