[1] Univ Auntonoma Barcelona, Inst Catala Paleontol Miquel Crusafont, Edifici ICTA ICP,C Columnes S-N,Campus UAB, Barcelona 08193, Spain
[2] Univ Kent, Sch Anthropol & Conservat, Marlowe Bldg, Canterbury CT2 7NR, England
[3] Univ Autonoma Barcelona, Campus UAB, Barcelona 08193, Spain
[4] Senckenberg Res Inst, Div Palaeoanthropol, Frankfurt, Germany
[5] Nat Hist Museum Frankfurt, Frankfurt, Germany
[6] Amer Museum Nat Hist, Div Anthropol, Cent Pk West & 79th St, New York, NY 10024 USA
[7] New York Consortium Evolutionary Primatol, New York, NY 10024 USA
[8] Kyoto Univ, Grad Sch Sci, Lab Phys Anthropol, Kyoto 6068502, Japan