[1] Univ Estadual Maranhao, Sao Luis, Brazil
[2] Univ Fed Maranhao, Sao Luis, Brazil
[3] Montpellier SupAgro, UMR Eco&Sols, CIRAD, INRA,IRD, Montpellier, France
[4] UMR iEES Paris, UPMC, CNRS, INRA,UPEC,IRD,AgroParisTech, Paris, France
[5] Univ Estadual Maranhao, PPGA Programa Posgrad Agroecol, Cidade Univ Paulo VI, Sao Luis, Maranhao, Brazil