[1] Univ Queensland, Sch Earth & Environm Sci, St Lucia, Qld 4067, Australia
[2] Univ Queensland, Sch Math & Phys, St Lucia, Qld 4067, Australia
[3] Commonwealth Sci & Ind Res Org CSIRO Oceans & Atmo, Queensland Biosci Precinct QBP, St Lucia, Qld 4067, Australia
[4] Univ New South Wales, Ctr Marine Sci & Innovat CMSI, Sydney, NSW 2052, Australia
[5] Sydney Inst Marine Sci, Mosman, NSW 2088, Australia
[6] Univ Libre Bruxelles ULB, Fac Sci, Dept Organism Biol, Syst Ecol & Resource Management, Ave FD Roosevelt 50, CPi 264-1, B-1050 Brussels, Belgium
[7] Vrije Univ Brussel VUB, Biol Dept, Lab Plant Biol & Nat Management, Ecol & Biodivers,VUB APNA WE, Pleinlaan 2, B-1050 Brussels, Belgium