[1] Univ Guadalajara, Herbario IBUG, Inst Bot, Dept Bot & Zool, Km 15-5 Guadalajara Nogales, Zapopan 45110, Jalisco, Mexico
[2] Univ Autonoma Occidente, Blvd Lola Beltran & Blvd Rolando Aijona,4 Marzo, Culiacan 80020, Sinaloa, Mexico
[3] Reserva Ecol Mineral Nuestra Senora Candelaria, Fac Biol, Cosala 80780, Sinaloa, Mexico
[4] Jardin Bot Benjamin F Johnston, Blvd Antonio Rosales 750 Sur, Los Mochis 81289, Sinaloa, Mexico
[5] Inst Tecnol & Estudios Super Occidente, Tlaquepaque 45604, Jalisco, Mexico