[1] Chinese Acad Sci, China Croatia Belt & Road Joint Lab Biodivers & Ec, CAS Key Lab Mt Ecol Restorat & Bioresource Utiliza, Beijing, Peoples R China
[2] Chinese Acad Sci, Chengdu Inst Biol, Ecol Restorat & Biodivers Conservat Key Lab Sichua, Beijing, Peoples R China
[3] Shanghai Normal Univ, Shanghai, Peoples R China
[4] Univ Copenhagen, Dept Biol, Terr Ecol Sect, Copenhagen, Denmark