[1] Univ Toulouse III Paul Sabatier, Lab Evolut & Divers Biol, UMR 5174, CNRS,IRD, 118 Route Narbonne, F-31062 Toulouse, France
[2] CNRS, Stn Ecol Theor & Expt, UAR 2029, Moulis, France
[3] Univ Exeter, Ctr Ecol & Conservat, Biosci, Penryn, Cornwall, England
[4] Univ Glasgow, Inst Biodivers Anim Hlth & Comparat Med, Coll Med Vet & Life Sci, Glasgow, Scotland
[5] Univ Liege, Freshwater & Ocean Sci Unit Res FOCUS, Lab Ecol & Conservat Amphibians LECA, B-4020 Liege, Belgium
[6] Univ Paris Saclay, U981, Gustave Roussy Canc Campus, INSERM, Villejuif, France
[7] Univ Ljubljana, Biotech Fac, Ljubljana, Slovenia