[1] Italian Inst Environm Protect & Res ISPRA, Area Protect Biodivers Habitats & Protected Marine, Via Vitaliano Brancati 60, I-00144 Rome, Italy
[2] Zool Stn Anton Dohrn SZN, Rome, Italy
[3] Univ Genoa, Dept Earth Environm & Life Sci DiSTAV, Genoa, Italy
[4] Natl Interuniv Consortium Marine Sci, Rome, Italy
[5] French Biodivers Agcy OFB, Marseille, France