[1] Mae Fah Luang Univ, Sch Sci, Chiang Rai 57100, Thailand
[2] Ubon Ratchathani Univ, Fac Sci, Dept Biol Sci, Ubon Ratchathani 34190, Thailand
[3] Chiang Mai Univ, Fac Sci, Dept Biol, Chiang Mai 50200, Thailand
[4] Chiang Mai Univ, Fac Sci, Res Ctr Microbial Divers & Sustainable Utilizat, Chiang Mai 50200, Thailand
[5] Acad Sci, Royal Soc Thailand, Bangkok, Thailand
[6] Meise Bot Garden, Nieuwelaan 38, B-1860 Meise, Belgium
[7] Federat Wallonie Bruxelles, Serv Gen Enseignement Super & Rech Sci, Brussels, Belgium