[1] Univ Fed Rio de Janeiro, Dept Vertebrados, Museu Nacl, Programa Posgrad Ciencias Biol Zool, Quinta Da Boa Vista, RJ, Brazil
[2] Univ Nacl Mayor San Marcos, Dept Ictiol, Museo Hist Nat, Lima, Peru
[3] Univ Nacl Mayor San Marcos, Fac Ciencias Biol, Museo Hist Nat, Lima, Peru
[4] Royal Ontario Museum, Dept Nat Hist, Toronto, ON, Canada
[5] Univ Toronto, Dept Ecol & Evolutionary Biol, Toronto, ON, Canada