[1] Res Dept, Meise Bot Garden, Nieuwelaan 38, B-1860 Meise, Belgium
[2] Univ Antwerp, Dept Biol ECOSPHERE, Univ Pl 1, B-2610 Antwerp, Belgium
[3] Sokoine Univ Agr SUA, Dept Crop Sci & Hort, POB 3005, Morogoro, Tanzania
[4] Luxembourg Inst Sci & Technol LIST, Environm Res & Innovat ERIN Dept, Observ Climate Environm & Biodivers OCEB, 41 rue Brill, L-4422 Belvaux, Luxembourg