[1] Northwest Univ, Coll Life Sci, Xian, Shaanxi, Peoples R China
[2] Univ Groningen, Groningen Inst Evolutionary Life Sci GELIFES, Groningen, Netherlands
[3] Univ Groningen, Groningen Inst Evolutionary Life Sci GELIFES, PO Box 11103, NL-9700 CC Groningen, Netherlands