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Sun Yat Sen Univ, Guangdong Prov Key Lab Plant Resources, Key Lab Gene Engn, Minist Educ,State Key Lab Biocontrol,Sch Life Sci, Guangzhou 510275, Guangdong, Peoples R China Sun Yat Sen Univ, Guangdong Prov Key Lab Plant Resources, Key Lab Gene Engn, Minist Educ,State Key Lab Biocontrol,Sch Life Sci, Guangzhou 510275, Guangdong, Peoples R China