[1] Univ Leon, Dept Ingn & Ciencias Agr, Area Fisiol Vegetal, Leon, Spain
[2] Univ Leon, Inst Biol Mol Genom & Prote INBIOMIC, Leon, Spain
[3] Univ Politecn Madrid UPM, Ctr Biotecnol & Genom Plantas, Inst Nacl Invest & Tecnol Agr & Alimentaria INIA C, Campus de Montegancedo UPM, Pozuelo De Alarcon, Madrid, Spain
[4] Escuela Tecn Super Ingn Agron Alimentaria & Biosis, Dept Biotecnol Biol Vegetal, Madrid, Spain