[1] Univ Calif Berkeley, Herbert Wertheim Sch Optometry & Vis Sci, Berkeley, CA 94720 USA
[2] Univ Calif Berkeley, Vis Sci Grad Program, Berkeley, CA 94720 USA
[3] Univ Rochester, Ctr Visual Sci, Rochester, NY 14627 USA
[4] Univ Rochester, Dept Biomed Engn, Rochester, NY USA
[5] Univ Rochester, Dept Neurosci, Rochester, NY USA
[6] Univ Rochester, Flaum Eye Inst, Dept Ophthalmol, Rochester, NY USA
[7] Univ Calif Berkeley, Helen Wills Neurosci Inst, Berkeley, CA 94720 USA