[1] Shandong Univ, Inst Ecol & Biodivers, Sch Life Sci, Qingdao 266237, Peoples R China
[2] Tsinghua Univ, Inst Environm & Ecol, Tsinghua Shenzhen Int Grad Sch, Shenzhen Key Lab Ecol Remediat & Carbon Sequestrat, Shenzhen 518055, Peoples R China
[3] Shandong Univ, Shandong Prov Engn & Technol Res Ctr Vegetat Ecol, Qingdao 266237, Peoples R China
[4] Shandong Univ, Qingdao Forest Ecol Res Stn, Natl Forestry & Grassland Adm, Qingdao 266237, Peoples R China
[5] Shandong Univ, Qingdao Key Lab Forest & Wetland Ecol, Qingdao 266237, Peoples R China
[6] Shandong Univ, Environm Res Inst, Qingdao 266237, Peoples R China
[7] Shandong First Med Univ, Dept Obstet & Gynecol, Shandong Key Lab Reprod Med, Shandong Prov Hosp, Jinan 250021, Shandong, Peoples R China