[1] KwaZulu Natal Museum, Dept Nat Sci, 237 Jabu Ndlovu St, ZA-3201 Pietermaritzburg, South Africa
[2] Rhodes Univ, Dept Zool & Entomol, POB 94, ZA-6140 Makhanda, South Africa
[3] Int Inst Trop Agr, Biodivers Ctr, 08 BP 0932 Tri Postal, Cotonou, Benin
[4] Royal Museum Cent Africa, Dept Biol, Leuvensesteenweg 13, B-3080 Tervuren, Belgium
[5] JEMU, Leuvensesteenweg 13, B-3080 Tervuren, Belgium