[1] INRAE, URFM, Avignon, France
[2] Avignon Univ, INRAE, UMR 1114, EMMAH, Avignon, France
[3] Ctr Recerca Ecol i Aplicac Forestals, CREAF, Bellaterra, Catalonia, Spain
[4] Montpellier Univ, CNRS, UMR 5175, CEFE,EPHE,IRD, Montpellier, France
[5] AgroParisTech, UPS, CNRS, UMR 8079A,ESE, Orsay, France