[1] Korea Res Inst Biosci & Biotechnol, Funct Biomat Res Ctr, Jeongeup 56212, South Korea
[2] Univ Sci & Technol UST, Dept Appl Biotechnol, Daejeon 34113, South Korea
[3] Chonnam Natl Univ, Coll Vet Med, Dept Vet Surg, Gwangju 61186, South Korea
[4] Chonnam Natl Univ, BK21 FOUR Program, Gwangju 61186, South Korea
[5] Chonnam Natl Univ, Dept Marine Bio Food Sci, Yeosu 59626, South Korea