[1] Qujing Normal Univ, Coll Biol Resource & Food Engn, Res Ctr Nat Hist & Culture, Ctr Yunnan Plateau Biol Resources Protect & Utiliz, Qujing 655011, Peoples R China
[2] Chiang Mai Univ, Fac Sci, Dept Biol, Chiang Mai 50200, Thailand
[3] Chiang Mai Univ, Ctr Excellence Microbial Divers & Sustainable Util, Chiang Mai 50200, Thailand
[4] King Saud Univ, Ctr Excellence Biotechnol Res CEBR, Riyadh, Saudi Arabia