[1] Univ Clermont Auvergne, INRAE, VetAgro Sup, UMR Herbivores, F-63122 St Genes Champanelle, France
[2] INRAE, Inst Agro, PEGASE, F-35590 St Gilles, France
[3] Univ Tours, INRAE, UMR BOA, F-37380 Nouzilly, France
[4] Univ Pau & Pays Adour, INRAE, NUMEA, F-64310 St Pee Sur Nivelle, France
[5] INRAE, DEPE, 147 Rue Univ, F-75338 Paris, France
[6] INRAE, LPGP, F-35000 Rennes, France
[7] IFREMER, Lab EM3B, Rue IIle Yeu, F-44311 Nantes, France
[8] Univ Toulouse, PSAE UR 1303, INRAE, ENVT, F-31076 Toulouse, France