[1] UMR CNRS 6553 Ecobio, Université de Rennes, 263 Avenue du Gal Leclerc, CS, Rennes cedex,74205, 35042, France
[2] Ecodiv USC INRAE 1499, University of Rouen Normandy, France
[3] Institute for Coastal and Marine Research, Nelson Mandela University, Port Elizabeth, South Africa