[1] CAS Key Laboratory of Coastal Environmental Processes and Ecological Remediation, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS), Shandong Key Laboratory of Coastal Environmental Processes, Yantai,264003, China
[2] Shandong Key Laboratory of Marine Ecological Restoration, Observation and Research Station of Laizhou Bay Marine Ecosystem, MNR, Shandong Marine Resources and Environment Research Institute, No. 216 Changjiang Road, Yantai,264006, China
[3] University of Chinese Academy of Sciences, Beijing,100049, China
[4] Key Laboratory of Ecological Restoration and Conservation of Coastal Wetlands in Universities of Shandong, The Institute for Advanced Study of Coastal Ecology, Ludong University, Yantai,264025, China
[5] School of Ocean, Yantai University, Yantai,264005, China