[1] Faculty of Fisheries, Kagoshima University, 4-50-20 Shimoarata, Kagoshima,890–0056, Japan
[2] Hatsukaichi Field Station, Fisheries Technology Institute, Japan Fisheries Research and Education Agency, 2-17-5 Maruishi, Hiroshima, Hatsukaichi,739-0452, Japan
[3] Yaeyama Field Station, Fisheries Technology Institute, Japan Fisheries Research and Education Agency, 148 Fukaiota, Ishigaki, Okinawa,907-0451, Japan
[4] Coastal and Inland Fisheries Ecosystems Division, Fisheries Technology Institute, Japan Fisheries Research and Education Agency, 1551-8 Tairamachi, Nagasaki,851-2213, Japan