[1] CSIRO Oceans & Atmosphere, Indian Ocean Marine Research Centre, University of Western Australia, M097, 35 Stirling Highway, Crawley,WA,6009, Australia
[2] Australian Research Council Centre of Excellence for Coral Reef Studies, James Cook University, Townsville,Queensland,4811, Australia
[3] CSIRO Oceans and Atmosphere, GPO Box 2583, Brisbane,Queensland,4001, Australia
[4] Marine Science Program, Biodiversity and Conservation Science, Department of Biodiversity, Conservation and Attractions, Kensington, W.A,6151, Australia
[5] Oceans Institute, University of Western Australia, Crawley,WA,6009, Australia