[1] Scripps Institution of Oceanography, University of California San Diego, 9500 Gilman Drive, La Jolla,CA,92093-0202, United States
[2] Faculty of Science and Engineering, Soka University, Hachioji, Tokyo,192-8577, Japan
[3] Wildlife Fish and Conservation Biology, University of California Davis, One Shields Avenue, Davis,CA,95616-8627, United States
[4] Faculty of Science, University of the Ryukyus, Nishihara,Okinawa,903-0213, Japan