[1] Univ Plymouth, Fac Sci & Engn, Sch Biol & Marine Sci, Plymouth PL4 8AA, Devon, England
[2] Univ Plymouth, Marine Inst, Plymouth PL4 8AA, Devon, England
[3] Univ Plymouth, Sustainable Earth Inst, Plymouth PL4 8AA, Devon, England
[4] Fed European Microbiol Soc, Delftechpk 37a, NL-2628 XJ Delft, Netherlands