[1] Staatl Museum Nat Kunde Stuttgart, Rosenstein 1, D-70191 Stuttgart, Germany
[2] Univ Hohenheim, Inst Biol, Fachgebiet Palaontol, D-70599 Stuttgart, Germany
[3] Friedrich Alexander Univ Erlangen Nurnberg FAU, Geozent Nordbayern, D-91054 Erlangen, Germany
[4] Univ Autonoma Barcelona, Inst Catala Paleontol Miquel Crusafont ICP CERCA, Edif ICTA ICP,C Columnes S-N,Campus UAB, Cerdanyola Del Valles 08193, Catalonia, Spain
[5] Univ Bristol, Sch Geog Sci, Univ Rd, Bristol BS8 1SS, England
[6] Chinese Acad Sci, Inst Tibetan Plateau Res, State Key Lab Tibetan Plateau Earth Syst Resources, Lincui Rd, Beijing 100101, Peoples R China