[1] Chinese Acad Sci, Kunming Inst Bot, Key Lab Plant Divers & Biogeog East Asia, Kunming 650201, Yunnan, Peoples R China
[2] Shandong Agr Univ, Coll Forestry, Tai An 271018, Shandong, Peoples R China
[3] Kunming Univ Sci & Technol, Fac Land Resource Engn, Kunming 650093, Yunnan, Peoples R China
[4] Chiba Univ, Grad Sch Hort, Chiba 2718510, Japan
[5] Chinese Acad Sci, Key Lab Trop Forest Ecol, Xishuangbanna Trop Bot Garden, Mengla 666303, Yunnan, Peoples R China