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Chinese Acad Sci, Inst Earth Environm, State Key Lab Loess & Quaternary Geol, Xian 710061, Peoples R China
Univ Chinese Acad Sci, Beijing 100049, Peoples R China
Univ Oxford, Recanati Kaplan Ctr, Dept Zool, Wildlife Conservat Res Unit, Tubney House, Tubney OX13 5QL, Oxon, England Chinese Acad Sci, Inst Earth Environm, State Key Lab Loess & Quaternary Geol, Xian 710061, Peoples R China