[1] Unity Environm Univ, New Gloucester, ME USA
[2] Oregon State Univ, Corvallis, OR 97331 USA
[3] Kyoto Univ, Kyoto, Japan
[4] Japan Soc Promot Sci, Kojimachi Business Ctr Bldg, Tokyo, Japan
[5] Azabu Univ, Dept Anim Sci & Biotechnol, Kanagawa, Japan
[6] Kyoto Univ, Wildlife Res Ctr, Kyoto, Japan
[7] Anicom Special Med Inst Inc, Res & Dev Sect, Yokohama, Japan
[8] Osaka Univ, Dept Comparat & Dev Psychol, Grad Sch Human Sci, Osaka, Japan