[1] Univ Douala Yabassi, Inst Sci Halieutiques, Dept Gest Ecosyst Aquat, POB 7236, Douala, Cameroon
[2] Leibniz Inst Evolut & Biodivers Sci, Museum Nat Kunde, Invalidenstr 43, D-10115 Berlin, Germany
[3] Cameroon Assoc Res Crustaceans & Other Aquat Resou, Unit Biodivers Discovery & Conservat, POB 43, Obala, Cameroon
[4] Nat Hist Museum, Dept Life Sci, London SW7 5BD, England
[5] Northern Michigan Univ, Dept Biol, Marquette, MI 49855 USA