[1] Inst Ecol AC, Red Biodivers & Sistemat, Xalapa 91073, Veracruz, Mexico
[2] Univ Autonoma Metropolitana Iztapalapa, Dept Biol, Div Ciencias Biol & Salud, Herbario Metropolitano, Mexico City 09310, Mexico
[3] Univ Guadalajara, Ctr Univ Ciencias Biol & Agr, Dept Bot & Zool, Zapopan, Jalisco, Mexico
[4] Inst Ecol AC, Carretera Antigua Coatepec 351,El Haya, Xalapa 91073, Veracruz, Mexico