[1] Chinese Acad Sci, Chendu Inst Biol, Chengdu 610213, Peoples R China
[2] Univ Chinese Acad Sci, Beijing 101408, Peoples R China
[3] Chinese Acad Sci, Chengdu Inst Biol, CAS Key Lab Mt Ecol Restorat & Bioresource Utiliza, Chengdu, Peoples R China
[4] Chinese Acad Sci, Chengdu Inst Biol, Biodivers Conservat Key Lab Sichuan Prov, Chengdu, Peoples R China
[5] CSIRO Agr & Food, Canberra, Australia