[1] Ohio State Univ, Dept Evolut Ecol & Organismal Biol, Aquat Ecol Lab, Columbus, OH 43212 USA
[2] Fisheries & Oceans Canada, Great Lakes Lab Fisheries & Aquat Sci, 867 Lakeshore Rd, Burlington, ON L7S IA1, Canada
[3] Univ Waterloo, Dept Earth & Environm Sci, Waterloo, ON N2L 3G1, Canada
[4] Univ Waterloo, Water Inst, Waterloo, ON N2L 3G1, Canada
[5] Heidelberg Univ, Dept Biol & Environm Sci, 310 East Market St, Tiffin, OH 44883 USA
[6] Heidelberg Univ, Natl Ctr Water Qual Res, 310 East Market St, Tiffin, OH 44883 USA
[7] Univ Toledo, Lake Erie Ctr, Dept Environm Sci, 2801 W Bancroft St, Toledo, OH 43606 USA