[1] Eotvos Lorand Univ, Dept Ethol Budapest, Pazmany Peter Setany 1-C, H-1117 Budapest, Hungary
[2] Hungarian Univ Agr & Life Sci, Inst Wildlife Management & Nat Conservat, Dept Wildlife Biol & Management, Pater Karoly utca 1, H-2100 Godollo, Hungary
[3] Budakesz Wildlife Pk, Hrsz 0210-12, H-2092 Budakeszi, Hungary