[1] Ocean Univ China, Coll Ocean & Atmospher Sci, Qingdao 266100, Peoples R China
[2] Shandong Marine Resource & Environm Res Inst, Shandong Prov Key Lab Restorat Marine Ecol, Yantai 264006, Peoples R China
[3] Ocean Univ China, Key Lab Marine Chem Theory & Technol, Minist Educ, Qingdao 266100, Peoples R China
[4] Minist Nat Resources, Inst Oceanog 4, Key Lab Trop Marine Ecosyst & Bioresource, Beihai 536000, Peoples R China