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Chinese Acad Sci, Northwest Inst Ecoenvironm & Resources, Gansu Gaolan Field Sci Observat & Res Stn Agr Eco, Lanzhou 730000, Gansu, Peoples R China Chinese Acad Sci, Northwest Inst Ecoenvironm & Resources, Gansu Gaolan Field Sci Observat & Res Stn Agr Eco, Lanzhou 730000, Gansu, Peoples R China