The relationships between modern pollen rain assemblages and vegetation from Sougna Mountain (Rif Mountains-Northern Morocco)

This study discovers the distribution of vegetation in the Sougna Mountain and its modern pollen representation and investigates the relationships with environment and land use variables. A total of 31 moss polsters were collected between 138 and 1364 m above sea-level from wooded to open areas (e.g. cork oak forest, pine reforestation, and shrubland). At each sampling site land use and environment variables were collected. The vegetation in the Sougna Mountain is scattered in a mosaic along three vegetation belts which are displayed in distinct pollen assemblages dominated by the pollen equivalents of the vegetation types (Quercus suber forest, matorrals of Erica spp. and Cistus spp. and Pinus pinaster reforestation). The variation between the variables and pollen data was performed using CCA analysis. The anthropogenic variables such as grazing, and cultivation, contribute to discrimination of the degraded and grazed areas. High precipitation and altitude have a rather good correlation to wooded communities and consequently a good relation to the tree cover. Our results show that the pollen types of Poaceae, Cannabis, Plantago, Nicotiana, Asteraceae, and Urtica are related mostly to human practice and distinguish the disrupted communities. The presence of those taxa in the pollen diagram must be interpreted carefully. A high taxonomic resolution is recommended to prevent the overestimation of the abundance of some taxa. Our results contribute to the advancement of palynological reconstructions of past vegetation and land use in Sougna Mountain and generally in the Rif landscape.