[1] Dept Environm Syst Sci, Ecosyst Management, ETH Zurich, CH-8092 Zurich, Switzerland
[2] Dept Civil Environm & Geomat Engn, Planning Landscape & Urban Syst, ETH Zurich, CH-8093 Zurich, Switzerland
[3] Inst Integrat Biol, Dept Environm Syst Sci, ETH Zurich, CH-8092 Zurich, Switzerland
[4] Ecosyst Management, Univ Str 16,CHN G72, CH-8092 Zurich, Switzerland