[1] Addis Ababa Univ, Ctr Environm Sci, King George VI St,POB 1176, Addis Ababa, Ethiopia
[2] Int Livestock Res Inst, Asmera Rd, Addis Ababa 5689, Ethiopia
[3] Univ Quebec Montreal, ESCER Ctr, 201 President Kennedy Ave, Montreal, PQ H2X 3Y7, Canada
[4] NORCE Norwegian Res Ctr, Bjerknes Ctr Climate Res, POB 22, N-5838 Bergen, Norway
[5] Addis Ababa Univ, IGSSA, King George VI St,POB 1176, Addis Ababa, Ethiopia