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Guangdong Acad Sci, Inst Zool, Guangdong Key Lab Anim Conservat & Resource Utiliz, Guangdong Publ Lib Wild Anim Conservat & Utilizat, 105 Xingang West Rd, Guangzhou 510260, Guangdong, Peoples R China Guangdong Acad Sci, Inst Zool, Guangdong Key Lab Anim Conservat & Resource Utiliz, Guangdong Publ Lib Wild Anim Conservat & Utilizat, 105 Xingang West Rd, Guangzhou 510260, Guangdong, Peoples R China